Why effort wins over pace and heart rate
Minutes-per-mile is the worst of the three on trail. The same effort can read as 9:00 on a flat road and 28:00 on a steep, rocky climb, so a pace target either pushes you to cook yourself on the hills or makes the flats feel like failure. Heart rate is better, but it drifts upward over the hours (cardiac drift), it lags on short climbs, and it spikes in heat, so a fixed heart-rate cap slowly lies to you as the day wears on.
Rate of perceived exertion (RPE), how hard it honestly feels on a 1 to 10 scale, is what most experienced ultrarunners actually pace by. And the plan is simple. Hold an easy, conversational effort (about RPE 3 to 4) for the first several hours, then let the terrain and your body set the pace. Keep the watch and the heart rate in view as cross-checks, but let effort be the boss.